Jonathan Moran

I am an ecologist with an interest in the interactions between plants, animals, and their environment. I have a PhD from the University of Aberdeen in the UK, and have been on the Faculty at Royal Roads since 2007. Current research topics include:

  1. Modelling of species distributions under projected climate change scenarios;
  2. Hummingbird migration monitoring in Western Canada;
  3. Investigation into ecological mismatch between hummingbirds and hummingbird-pollinated plants; and 
  4. The use of metals as intrinsic markers for habitat use by birds.

Selected Publications

Journal Articles

Moran, J.A., Gray, L.K., Clarke, C. & Chin, L. (2013) Capture mechanism in palaeotropical pitcher plants (Nepenthaceae) is constrained by climate. Annals of Botany doi: 10.1093/aob/mct195 

Moran, J.A., Finlay, J.C., Wassenaar, L., Isaac, L.A., Hutcheson, C. & Wethington, S. (2012) An exploration of migratory connectivity of the Rufous Hummingbird, Selasphorus rufus, using feather deuterium. Journalof Ornithology 154: 423-430.

Moran, J.A., Clarke, C., Greenwood, M., Chin, L. (2012) Tuning of colour contrast signals to visual sensitivity maxima of tree shrews by three Bornean highland Nepenthes species. Plant Signaling & Behavior 7: 1267-1270.

Moran, J.A., Clarke, C., Gowen, B.E. (2012) The use of light in prey capture by the tropical pitcher plant Nepenthes aristolochioides. Plant Signaling & Behavior 7: 957-960.

Moran, J.A. (2012) Nepenthes baramensis C. Clarke, J. A. Moran & C. C. Lee, sp. nov. AIPC Special Issue 4: 18.

Bazile, V., Moran, J.A., le Moguedec, G., Marshall, D.G., Gaume, L. (2012) A carnivorous plant fed by its ant symbiont: a unique multi-faceted nutritional mutualism. PLoS ONE 7: e36179. doi:10.1371/journal.pone.0036179.

Clarke, C. & Moran, J.A. (2011) Incorporating ecological context: a revised protocol for the preservation of Nepenthes pitcher plant specimens (Nepenthaceae). Blumea 56: 225-228.

Clarke, C.M., Moran, J.A. & Lee, C.C. (2011) Nepenthes baramensis (Nepenthaceae) - a new species from Northwestern Borneo. Blumea 56: 229-233.

Moran, J.A., Hawkins, B.J., Gowan, B. & Turner, S. (2010) Ion fluxes across the pitcher walls of three Bornean Nepenthes pitcher plant species: flux rates and gland distribution patterns reflect nitrogen sequestration strategies. Journal of Experimental Botany 61: 1365-1374.

Moran, J.A. & Clarke, C.M. (2010) Invited Review: The carnivorous syndrome in Nepenthes pitcher plants: current state of knowledge and potential future directions. Plant Signaling & Behavior 5 (6): 1-5.

Clarke, C., Moran, J.A. & Chin, L. (2010) Mutualism between tree shrews and pitcher plants- perspectives and avenues for future research. Plant Signaling & Behavior 5 (10): 1-3.

Chin, L., Moran, J.A. & Clarke, C. (2010) Trap geometry in three giant montane pitcher plant species from Borneo is a function of tree shrew body size. New Phytologist 186: 461-470.

Hutcheson, C., Hendrix, L. & Moran, J.A. (2010) An isotopic analysis of migratory connectivity in ruby-throated hummingbirds (Archilochus colubris). North American Bird Bander Jan-Mar 2010: 5-11.

Clarke, C.M., Bauer, U., Lee, C.C., Tuen, A.A., Rembold, K. & Moran, J.A. (2009). Tree shrew lavatories: a novel nitrogen sequestration strategy in a tropical pitcher plant. Biology Letters 5: 632-635. 

Moran, J.A. (2006) Life and death in a pitcher. Natural History October 2006: 56-62. 

Ethier, G.J., Livingston, N.J., Harrison, D.L., Black, T.A. & Moran, J.A. (2006) Low stomatal and internal conductance to CO2 vs. Rubisco deactivation as determinants of the photosynthetic decline of aging evergreen leaves. Plant, Cell, and Environment 29: 2168-2184. 

Moran, J.A., Clarke, C.M. & Hawkins, B.J. (2003) From carnivore to detritivore? Isotopic evidence for leaf litter utilization by the tropical pitcher plant Nepenthes ampullaria. International Journal of Plant Sciences 164: 635-639. 

Hawkins, B.J. & Moran J.A. (2003) Growth responses of Abies amabilis advance regeneration to overstory removal, nitrogen fertilization and release from Vaccinium competition. Forest Science 49: 799-806. 

Brang, P., Moran, J.A., Puttonen, P. & Vyse, A. (2003) Regeneration of Picea engelmannii and Abies lasiocarpa in high-elevation forests of south-central British Columbia depends on nurse logs. Forestry Chronicle 79: 273-279. 

Mitchell, A.K., Bown, T., Dunsworth, B.G. & Moran, J.A. (2003) Above-ground biomass predicts growth limitation in amabilis fir and western hemlock. Forestry Chronicle 79: 285-290. 

Moran, J.A., Merbach, M.A., Livingston, N.J., Clarke, C.M. & Booth, W.E. (2001) Termite prey specialization in the pitcher plant Nepenthes albomarginata - evidence from stable isotope analysis. Annals of Botany 88: 307-311. 

Percy, R.E., Livingston, N.J., Moran, J.A. & Von Aderkas, P. (2001) Desiccation, cryopreservation and water relations parameters of white spruce (Picea glauca) and interior spruce (Picea glauca x engelmannii complex) somatic embryos. Tree Physiology 21: 1303-1310. 

Moran, J.A., Barker, M.G., Moran, A.J., Becker, P. & Ross, S.M. (2000) A comparison of the soil water, nutrient status and litterfall characteristics of tropical heath and mixed dipterocarp forest sites in Brunei. Biotropica 32: 2-13. 

Moran, J.A., Mitchell, A., Goodmanson, G. & Stockburger, K. (2000) Differentiation among effects of nitrogen fertilisation treatments of conifer seedlings by foliar reflectance: a comparison of methods. Tree Physiology 20: 1113-1120. 

Moran, J.A., Booth, W.E. & Charles, J.K. (1999) Aspects of pitcher morphology and spectral characteristics of six Bornean Nepenthes pitcher plant species: Implications for prey capture. Annals of Botany 83: 521-528. 

Moran, J.A. & Moran, A.J. (1998) Foliar reflectance and vector analysis reveal nutrient stress in prey-deprived pitcher plants (Nepenthes rafflesiana). International Journal of Plant Sciences 159: 996-1001. 

Moran, J.A. (1996) Pitcher dimorphism, prey composition and the mechanisms of prey attraction in the pitcher plant Nepenthes rafflesiana in Borneo. Journal of Ecology 84: 515-525. 

Deans, J.D., Moran. J.A. & Grace, J. (1996) Biomass relationships for tree species in regenerating semi-deciduous tropical moist forest in Cameroon. Forest Ecology and Management 88: 215-225.

Clarke, C.M. & Moran, J.A. (1994) A further record of aerial pitchers produced by Nepenthes ampullaria (Jack). Malayan Nature Journal 47: 321-323. 

Moran, J.A. (1993) Visitors to the flowers of the pitcher plant Nepenthes rafflesiana. Brunei Museum Journal 8: 73-75.

Moran, J.A. (1993) Pitcher allocation strategy of the pitcher plant Nepenthes rafflesiana in Brunei. Brunei Museum Journal 8: 77-80.

Moran, J.A. (1993) The effect of pitcher wing removal on prey capture by the pitcher plant Nepenthes rafflesiana. Brunei Museum Journal 8: 81-82.

Moran, J.A. (1993) Misumenops nepenthicola: The top predator of the Nepenthes food web? Brunei Museum Journal 8: 83-84.

Book Chapters/Contributions

Clarke, C.M. & Moran, J.A. (2001) Chapter 2. The Ecology of Nepenthes. In: Clarke, C.M. Nepenthes of Sumatra and Peninsular Malaysia. Natural History Publications, Kota Kinabalu, Malaysia. pp 29-75. 

Moran, J.A. (1997) Catching Prey. In: Clarke, C.M. Nepenthes of Borneo. Natural History Publications, Kota Kinabalu, Malaysia. pp 32-35, 39. 

Moran, J.A. (1994) Pitcher Plants. In: Cranbrook, Earl of & Edwards, D.S. Belalong: A Tropical Rainforest. Sun Tree Publishing, Singapore. pp 171-173. 

Conference/Symposium Proceedings

Moran, J.A. & Clarke, C.M. (2010) A review of the nutrient sequestration strategies of some Bornean Nepenthes species. Invited paper, Sarawak Nepenthes Summit, Kuching, Malaysia, 18-21 August 2007. pp 8-18. 

Reimchen, T.E., Mathewson, D., Hocking, M., Moran, J.A. & Harris, D. (2003) Isotopic evidence for enrichment of salmon-derived nutrients in vegetation, soil and insects in riparian zones in coastal British Columbia. In: Stock, J.G. (ed.) Nutrients in Salmonid Ecosystems: Sustaining Production and Biodiversity. American Fisheries Society Symposium 34: 59-69. 

Conference/Symposium Presentations

A Stable Isotope Study of Migratory Connectivity in Rufous Hummingbirds. Presentation to the Hummingbird Conservation Symposium / Cooper Ornithological Society 79th Meeting, Tucson, Arizona, 17-18th April 2009.

A Review of the Nutrient Sequestration Strategies of Some Bornean Nepenthes Species. Invited Speaker, Sarawak Nepenthes Summit, Kuching, Malaysia, 18-21 August 2007. 

Selasphorus rufus: Results from 10 Years of Banding Data from British Columbia, Canada. Presentation to “Hummingbird Monitoring and Conservation along North American Migratory Corridors: A Workshop in Memory of Bill Calder”, University of Guadalajara, Mexico, 22-29th February 2007.